Growth of the pollen tube of angiosperms is an important model study of the functioning of the information system in plants. Over the last 10 Anniversary cultured in vitro pollen tubes have been widely used in biochemical and cytofizjologicznych. The research focused on understanding the molecular events which added calcium ions in the mechanisms responsible for the regulation of growth and orientation of the pollen tube. In the present work presents the results of research on the regulation of calcium homeostasis in łagiewkach pollen, functioning fosfatydyloinozytolowego signal transduction pathway during its growth and the role of cytosolic free Ca 2 + levels in the reorientation of pollen tube growth direction.